What is the relationship between seive cells and companion cells in phloem tissue?

Each sieve tube element is normally associated with one or more nucleated companion cells, to which they are connected by plasnodesmata (channels between the cells). Each companion cell is derived from the same mother cell as its associated sieve tube member. Sieve tube members have no cell nucleus, ribosomes, or vacues. Thus, they depend on companion cells to provide proteins, ATP, and signalling molecules