What class of words in Modern English retains grammatical gender?

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The only traces of the Old English gender system are found in the system of pronoun-antecedent agreement, although this is now based on the sex, gender identity, or perceived sexual characteristics, of the pronoun's referent. Another manifestation of natural gender that continues to function in English is the use of certain nouns to refer specifically to persons or animals of a particular sex: widow/widower, actor/actress, etc.