Which of the following does natural selection act directly upon? Alleles, genes, individual organisms, or mutations?

Natural selection act directly upon Alleles

Explanation:

Genes, such as that of fur color, have many variants called alleles. Natural selection act on the alleles and chooses the best fit for an environment and eliminates the disadvantageous ones from the population over generations. Mutations are a rare occurrence that increases the number of allelic variants of a genes on which natural selection can act upon.