What connects the upper motor neurons to lower motor neurons?

The upper motor neurons synapse in the spinal cord connect with anterior horn cells of lower motor neurons, usually via interneurons. The anterior horn cells are the cell bodies of the lower motor neurons and are located in the grey matter of the spinal cord.

Explanation:

Interneurons are the central nodes of neural circuits, enabling communication between the upper motor neurons, sensory or motor neurons located in the brain and spinal cord and they send signals to lower motor neurons or central nervous system (CNS) in the brain stem and spinal cord. When they get a signal from the upper motor neurons, they send another signal to your muscles to make them contract. They play vital roles in reflexes, neuronal oscillations, and neurogenesis in the adult mammalian brain.

Renshaw cells are among the very first identified interneurons. They are excited by the axon collaterals of the motor neurons. In addition, Renshaw cells make inhibitory connections to several groups of motor neurons.